AskDefine | Define stipule

Dictionary Definition

stipule n : a small leafy outgrowth at the base of a leaf or its stalk; usually occurring in pairs and soon shed

User Contributed Dictionary



Latin for "stubble"


IPA: /ˈstɪpjuːl/


  1. Basal appendage of a typical leaf of a flowering plant, usually appearing paired beside the petiole although sometimes absent or highly modified.

Derived terms

Extensive Definition

In botany, stipule (Latin stipula: straw, stalk) is a term coined by Linnaeus which refers to outgrowths borne on either side of the base of a leafstalk (the petiole). A pair of stipules is considered part of the anatomy of the leaf of a typical flowering plant, although in many species the stipules are inconspicuous or entirely absent (and the leaf is then termed exstipulate).
Stipules are morphologically variable and might appear as glands, scales, hairs, spines, or laminar (leaf-like) structures. A relationship exists between the anatomy of the stem node and the presence or absence of stipules. Most plants with trilacunar nodes have stipules; species with unilacunar nodes lack stipules (Sinnott and Bailey, 1914, cited in Esau, 1953).



General references

  • Esau, K. 1953. Plant Anatomy. Second Edition. John Wiley & Sons, Inc., New York, London, Sidney. 767 pp.
  • Sinnott, E. W. and I. W. Bailey. 1914. Investigations on the phylogeny of the angiosperms. 3. Nodal anatomy and the morphology of stipules. Amer. J. Bot.,1: 441-453.

External links

stipule in Catalan: Estípula
stipule in Czech: Palist
stipule in German: Nebenblatt
stipule in Spanish: Estípula
stipule in French: Stipule
stipule in Italian: Stipola
stipule in Dutch: Steunblaadje
stipule in Polish: Przylistek
stipule in Portuguese: Estípula
stipule in Serbian: Лисни залисци
stipule in Serbo-Croatian: Lisni zalisci
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